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October 10, 2013 9:13pm CST
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Propagules. In general, there are four major classes of hyperparasites: obligate bacterial pathogens, hypoviruses, facultative parasites, and predators. Pasteuria penetrans is an obligate bacterial pathogen of rootknot nematodes that has been used as a BCA. Hypoviruses are hyperparasites. A classical example is the virus that infects Cryphonectria parasitica, a fungus causing chestnut blight, which causes hypovirulence, a reduction in diseaseproducing capacity of the pathogen. The phenomenon has controlled the chestnut blight in many places (Milgroom and Cortesi 2004). However, the interaction of virus, fungus, tree, and environment determines the success or failure of hypovirulence. Pythium oligandrum). And, a single fungal pathogen can be attacked by multiple hyperparasites. For example, Acremonium alternatum, Acrodontium crateriforme, Ampelomyces quisqualis, Cladosporium oxysporum, and Gliocladium virens are just a few of the fungi that have the capacity to parasitize powdery mildew pathogens (Kiss 2003). Paecilomyces lilacinus and Dactylella oviparasitica). In contrast to hyperparasitism, microbial predation is more general and pathogen nonspecific and generally provides less predictable levels of disease control. Some BCAs exhibit predatory behavior under nutrientlimited conditions. However, such activity generally is not expressed under typical growing conditions. For example, some species of Trichoderma produce a range of enzymes that are directed against cell walls of fungi. However, when fresh bark is used in composts, Trichoderma spp. do not directly attack the plant pathogen, Rhizoctonia solani. But in decomposing bark, the concentration of readily available cellulose decreases and this activates the chitinase genes of Trichoderma spp., which in turn produce chitinase to parasitize R. solani (Benhamou and Chet 1997). Antibioticmediated suppression Antibiotics are microbial toxins that can, at low concentrations, poison or kill other microorganisms. Most microbes produce and secrete one or more compounds with antibiotic activity. In some instances, antibiotics produced by microorganisms have been shown to be particularly effective at suppressing plant pathogens and the diseases they cause.
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